This article takes a lot of words to state what is, ultimately, common sense for the already-enlightened. That is often what research articles do. At least it spells its findings out in such obvious, easy-to-follow reasoning that the reader is left to regard the conclusion as obviously correct.

As mentioned by others already, it totally ignores the important role of anti-queer religious indoctrination. But I am at least somewhat convinced that religion, though a highly effective tool for reenforcing prejudice and hate, may not actually be the ultimate source of motivation for such prejudice. That source, may indeed be as the author suggests, our culture's reverence for toxic masculinity values and its emphasis on rigidly segregated gender roles. Religion itself becomes shaped by those cultural values to be used as a reenforcing "weapon" in that campaign. Chicken, meet egg.